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The statistical analysis of phylogenetic footprints in the two known horn shark Hox clusters and the four mammalian clusters shows that the shark HoxN cluster is HoxD-like. This finding implies that the most recent common ancestor of jawed vertebrates had at least four Hox clusters, including those which are orthologous to the four mammalian Hox clusters.


Sonja J Prohaska, Claudia Fried, Chris T Amemiya, Frank H Ruddle, Günter P Wagner, Peter F Stadler. The shark HoxN cluster is homologous to the human HoxD cluster. Journal of molecular evolution. 2004 Feb;58(2):212-7

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PMID: 15042342

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